diff --git a/spring-data-mongodb/src/main/java/org/springframework/data/mongodb/core/EntityOperations.java b/spring-data-mongodb/src/main/java/org/springframework/data/mongodb/core/EntityOperations.java
index bb2d2ca5a6..db07166b5a 100644
--- a/spring-data-mongodb/src/main/java/org/springframework/data/mongodb/core/EntityOperations.java
+++ b/spring-data-mongodb/src/main/java/org/springframework/data/mongodb/core/EntityOperations.java
@@ -283,6 +283,10 @@ public <T> TypedOperations<T> forType(@Nullable Class<T> entityClass) {
 	 * @see EntityProjectionIntrospector#introspect(Class, Class)
 	 */
 	public <M, D> EntityProjection<M, D> introspectProjection(Class<M> resultType, Class<D> entityType) {
+
+		if (!queryMapper.getMappingContext().hasPersistentEntityFor(entityType)) {
+			return (EntityProjection) EntityProjection.nonProjecting(resultType);
+		}
 		return introspector.introspect(resultType, entityType);
 	}
 
diff --git a/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/MongoTemplateTests.java b/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/MongoTemplateTests.java
index 30978bce96..c93c8ade66 100644
--- a/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/MongoTemplateTests.java
+++ b/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/MongoTemplateTests.java
@@ -2526,6 +2526,26 @@ public void findAndReplaceShouldProjectReturnedObjectCorrectly() {
 		assertThat(projection.getName()).isEqualTo("Walter");
 	}
 
+	@Test // GH-4300
+	public void findAndReplaceShouldAllowNativeDomainTypesAndReturnAProjection() {
+
+		MyPerson person = new MyPerson("Walter");
+		person.address = new Address("TX", "Austin");
+		template.save(person);
+
+		MyPerson previous = template.findAndReplace(query(where("name").is("Walter")),
+				new org.bson.Document("name", "Heisenberg"), FindAndReplaceOptions.options(), org.bson.Document.class,
+				"myPerson", MyPerson.class);
+
+		assertThat(previous).isNotNull();
+		assertThat(previous.getAddress()).isEqualTo(person.address);
+
+		org.bson.Document loaded = template.execute(MyPerson.class, collection -> {
+			return collection.find(new org.bson.Document("name", "Heisenberg")).first();
+		});
+		assertThat(loaded.get("_id")).isEqualTo(new ObjectId(person.id));
+	}
+
 	@Test // DATAMONGO-407
 	public void updatesShouldRetainTypeInformationEvenForCollections() {
 
diff --git a/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/MongoTemplateUnitTests.java b/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/MongoTemplateUnitTests.java
index 3f4a63232e..6ae652ef85 100644
--- a/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/MongoTemplateUnitTests.java
+++ b/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/MongoTemplateUnitTests.java
@@ -2392,6 +2392,17 @@ void templatePassesOnTimeSeriesOptionsWhenNoTypeGiven() {
 				.isEqualTo(new com.mongodb.client.model.TimeSeriesOptions("time_stamp").toString());
 	}
 
+	@Test // GH-4300
+	void findAndReplaceAllowsDocumentSourceType() {
+
+		template.findAndReplace(new Query(), new Document("spring", "data"), FindAndReplaceOptions.options().upsert(),
+				Document.class, "coll-1", Person.class);
+
+		verify(db).getCollection(eq("coll-1"), eq(Document.class));
+		verify(collection).findOneAndReplace((Bson) any(Bson.class), eq(new Document("spring", "data")),
+				any(FindOneAndReplaceOptions.class));
+	}
+
 	class AutogenerateableId {
 
 		@Id BigInteger id;
diff --git a/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/ReactiveMongoTemplateTests.java b/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/ReactiveMongoTemplateTests.java
index 6ca3808c9a..7740b5a7cf 100644
--- a/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/ReactiveMongoTemplateTests.java
+++ b/spring-data-mongodb/src/test/java/org/springframework/data/mongodb/core/ReactiveMongoTemplateTests.java
@@ -729,6 +729,32 @@ void findAndReplaceShouldProjectReturnedObjectCorrectly() {
 				}).verifyComplete();
 	}
 
+	@Test // GH-4300
+	public void findAndReplaceShouldAllowNativeDomainTypesAndReturnAProjection() {
+
+		MongoTemplateTests.MyPerson person = new MongoTemplateTests.MyPerson("Walter");
+		person.address = new Address("TX", "Austin");
+		template.save(person) //
+				.as(StepVerifier::create) //
+				.expectNextCount(1) //
+				.verifyComplete();
+
+		template
+				.findAndReplace(query(where("name").is("Walter")), new org.bson.Document("name", "Heisenberg"),
+						FindAndReplaceOptions.options(), org.bson.Document.class, "myPerson", MongoTemplateTests.MyPerson.class)
+				.as(StepVerifier::create) //
+				.consumeNextWith(actual -> {
+					assertThat(actual.getAddress()).isEqualTo(person.address);
+				}).verifyComplete();
+
+		template.execute(MongoTemplateTests.MyPerson.class, collection -> {
+			return collection.find(new org.bson.Document("name", "Heisenberg")).first();
+		}).as(StepVerifier::create) //
+				.consumeNextWith(loaded -> {
+					assertThat(loaded.get("_id")).isEqualTo(new ObjectId(person.id));
+				}).verifyComplete();
+	}
+
 	@Test // DATAMONGO-1827
 	void findAndReplaceShouldReplaceObjectReturingNew() {